Sent into bat, Dhawan (94) and captain Virat Kohli (48) led India's recovery in a 97-run second-wicket partnership, before Mendis, more than three years after his mystery was decoded by the Indian batsmen, came back to trouble them with figures of 4/60 at the Khan Shaheb Osman Ali Stadium.
Mendis castled Kohli and Dhawan with his carrom ball while off-spinner Sachitra Senanayake scalped 3/41 as India's new-look middle order sorely missed Mahendra Singh Dhoni.
Thanks to the duo's exploits, India slipped after being comfortably placed at 175 for two in the 36th over.
Ajinkya Rahane (22), Ambati Rayudu (18) and Dinesh Karthik (4) looked good but could not covert their starts, while Stuart Binny did not trouble the scorers after being out for a zero.
Having come into the limelight with his career-best 6/13 against India in Karachi in 2008, Mendis got a reality check after the currently out-of-favour Virender Sehwag unleashed his fury on him the next year.
But that was then and on this day Mendis, who was included in place of medium pacer Suranga Kamal, did not let the Lankans down.
The pitch behaved differently with the ball staying low right from the very first over. It was the perfect condition to test the openers who were in the middle of a prolonged lean patch.
Sharma, particularly, struggled against the odd ball as Sri Lanka, who were playing with three specialist spinners, brought on offie Senanayake in the sixth over.
Lanka seemed in perfect control, conceding only 37 runs in the first power play which also yielded Sharma's wicket.
After all his struggle, Sharma'a anxious stay finally came to an end when he got to out to a dubious decision. Umpire Nigel Llong ruled the batsman out despite being long way down the track.
The breakthrough incidentally turned around India's fortunes with last match centurion Kohli starting off with a boundary.
At the other end, Dhawan looked to have got his fluency back and Kohli's reassuring presence further calmed the left-hander's nerves.
Sri Lanka's pace attack lacked bite after skipper Angelo Mathews left the field midway into his third over with a niggle as the Lankans resorted to an all-spin attack with Mendis coming into bowl in the 16th over.
With Kohli in exquisite form and Dhawan back among runs, the duo looked in control of the proceedings.
Dhawan completed his half-century in 68 balls, the left-hander's first 50-plus score in nine matches. Kohli punished Chaturanaga de Silva before Dhawan joined the party with a boundary and a six against Perera, as India raced to 128/1 in 26 overs, when a smart decision to bring in Mendis from the pavilion end did the trick.
Mendis, who had a listless 4-0-22-0 in the first spell, bowled a carrom ball to castle the in-form Kohli, who fell two short of a half-century.
The prized scalp of Kohli broke India's back and the new- look middle order, without Dhoni, could never recover.
India took the batting power play early in the 33rd over but the inexperienced middle order in Rahane, Rayudu, Karthik and Binny failed to cash in.
Only 28 runs and Rahane's wicket in the batting powerplay was the beginning of India's downfall.
Mendis two wickets in one over, including that of Dhawan with another carrom ball, to deny the lefthander a century.
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